Citations: Minkina, Ł.; Anichtchenko, A.; Vasiljeva, A.; Skelley, P. E. 2022: Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, Daugavpils 22 (1), S. 81-101, 12 389, pp. 82